Ponderosa and lodgepole pines and some spruce and fir species Colorado shed older, interior needles as part of an annual growth cycle in September and October.
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Columbia Gorge
My Conifers Are Losing Needles! Should I Be Concerned? - Independent Tree
Best National Parks to Visit in October & Fall Colors
Hydrological effects of forest transpiration loss in bark beetle-impacted watersheds
5 of the Most Common Tree Diseases of Wisconsin (And What You Can Do) Earth Development
Category: Trees - K-State Research and Extension Horticulture Newsletter
Types of Coniferous Trees: Forest Conifers Vs. Deciduous Vs. Evergreen
JeffCo Master Gardeners: Ponderosa pines, spruce may exhibit 'winter burn' by Colorado State Forest Service
Meet the beetles
Help! I think my Ponderosa Pine is Dying!
Why Do Pine Trees Lose Their Needles
Borers of Pines and Other Needle Bearing Evergreens in Landscapes
Pinus albicaulis